解释：句中有两组插入语，每组两个插入语，尤其是第二组，把rate of从中间劈开。不过因为句义尚好理解，所以本句只是有一点别扭，但不算太难。
意群训练：In the periods of peak zooplankton abundance,that is,in the late spring and in the summer,Haney recorded maximum daily community grazing rates,for nutrient-poor lakes and bog lakes，respectively,of 6.6 percent and 114 percent of daily phytoplankton production.
